Prof. Dr. Henrik Krehenwinkel

Publications:

  1. Krehenwinkel, H., Junk, I., Hans, J., Perez-Lamarque, B., Weber, S., Stothut, M., ... & Morlon, H. (2025). Archived natural DNA samplers reveal four decades of biodiversity change across the tree of life. Nature Ecology and Evolution.
  2. Schmidt, A., Schilbach, L., Schanowski, A., Grevé, M. E., Ulrich, C., Baden, C. M., & Krehenwinkel, H. (2025). Flower-derived environmental DNA reveals community diversity, species abundances and ecological interactions in bee pollinators. Environmental DNA.
  3. Hans, J., Stothut, M., Schubert, C., Gold, E., Chung, D., Lee, J., ... & Krehenwinkel, H. (2025). eDNA metabarcoding of archived leaf samples reveals arthropod diversity decline in South Korean but not in German forest ecosystems. Insect Conservation and Diversity.
  4. Nariman, N., Entling, M. H., Krehenwinkel, H., & Kennedy, S. (2025). The Microbiome of an Invasive Spider: Reduced Bacterial Richness, but no Indication of Microbial-Mediated Dispersal Behaviour. Microbial Ecology, 88(1), 70.
  5. Plewnia, A., Krehenwinkel, H., & Heine, C. (2025). Towards low-cost and PCR free field-based community metabarcoding. Methods in Ecology and Evolution.
  6. Schöneberg, Y., Audisio, T. L., Ben Hamadou, A., Forman, M., Král, J., Kořínková, T., ... & Kennedy, S. (2025). Three Novel Spider Genomes Unveil Spidroin Diversification and Hox Cluster Architecture: Ryuthela nishihirai (Liphistiidae), Uloborus plumipes (Uloboridae) and Cheiracanthium punctorium (Cheiracanthiidae). Molecular Ecology Resources, 25(1), e14038.
  7. Stothut, M., Mahla, L., & Krehenwinkel, H. (2025). Insektenvielfalt im Wandel der Zeit–neue Erkenntnisse aus Herbarien. BIOspektrum, 31(3), 262-265.
  8. Akhmedov, A., Bobokandov, N., Krehenwinkel, H., Rzepecki, A., Klein, T., Villar, J. L., & Thomas, F. M. (2025). Wood Anatomical and Hydraulic Traits of Tamarix Species Across a Large Eurasian Gradient Show a Stronger Climatic Than Phylogenetic Signal. Journal of Biogeography.
  9. Melcher, A. C., Krehenwinkel, H., Harms, D., & Birkhofer, K. (2025). Estimating Required Sample Sizes for Gut Content Metabarcoding Studies of Dietary Diversity in Spiders. Journal of Applied Entomology.
  10. Lötters, S., Böning, P., Bailon, S., Castañeda, J. D. B., Boistel, R., Catenazzi, A., ... & Plewnia, A. (2025). A roadmap for harlequin frog systematics, with a partial revision of Amazonian species related to Atelopus spumarius. Zootaxa, 5571(1), 1-76.
  11. Rhee, H., Naber, S., Krehenwinkel, H., & Hochkirch, A. (2025). Wildfires induce a reduction in body size and morphological variation of an insular endemic insect. Ecological Entomology.
  12. Steigerwald, E., Paetsch, J., Drück, D., Fritsch, J., Klaka, M., Knope, M. L., ... & Krehenwinkel, H. (2025). Of Islands on Islands: Natural Habitat Fragmentation Drives Microallopatric Differentiation in the Context of Distinct Biological Assemblages. Environmental DNA, 7(2), e70091.
  13. Stothut, M., Mahla, L., Backes, L., Weber, S., Avazzadeh, A., Moradmand, M., & Krehenwinkel, H. (2024). Recovering plant-associated arthropod communities by eDNA metabarcoding historical herbarium specimens. Current Biology, 34(18), 4318-4324.
  14. Stothut, M., Kühne, D., Ströbele, V., Mahla, L., Künzel, S., & Krehenwinkel, H. (2024). Environmental DNA metabarcoding reliably recovers arthropod interactions which are frequently observed by video recordings of flowers. Environmental DNA, 6(3), e550.
  15. Wolff, J. O., Kennedy, S. R., Houghton, M., Pascoe, P., Gajski, D., Derkarabetian, S., ... & Renault, D. (2024). Infrequent long-range dispersal and evolution of a top terrestrial arthropod predator in the sub-Antarctic. The American Naturalist, 204(2), 191-199.
  16. Fuhrmann, N., Brasseur, M. V., Bakowski, C. E., Podsiadlowski, L., Prost, S., Krehenwinkel, H., & Mayer, C. (2024). Chromosome-level genome assembly of the viviparous eelpout Zoarces viviparus. Genome Biology and Evolution, 16(8), evae155.
  17. Melcher, A. C., Weber, S., Birkhofer, K., Harms, D., & Krehenwinkel, H. (2024). To pool or not to pool: Pooled metabarcoding does not affect estimates of prey diversity in spider gut content analysis. Ecological Entomology, 49(6), 768-778.
  18. Friedrich, S., De Vivo, M., Baden, C. U., Grevé, M. E., Maus, C., Udelhoven, T., & Krehenwinkel, H. (2024). The quantity of deposited environmental DNA in plant–arthropod interactions depends on taxon, abundance and interaction time. Ecological Entomology, 49(6), 989-993.
  19. Weber, S., Stothut, M., Mahla, L., Kripp, A., Hirschler, L., Lenz, N., ... & Krehenwinkel, H. (2024). Plant‐derived environmental DNA complements diversity estimates from traditional arthropod monitoring methods but outperforms them detecting plant–arthropod interactions. Molecular Ecology Resources, 24(2), e13900.
  20. Gajski, D., Wolff, J. O., Melcher, A., Weber, S., Prost, S., Krehenwinkel, H., & Kennedy, S. R. (2024). Facilitating taxonomy and phylogenetics: An informative and cost-effective protocol integrating long amplicon PCRs and third-generation sequencing. Molecular Phylogenetics and Evolution, 192, 107988.
  21. Junk, I., Schmitt, N., & Krehenwinkel, H. (2023). Tracking climate-change-induced biological invasions by metabarcoding archived natural eDNA samplers. Current Biology, 33(18), R943-R944.
  22. Graham, N. R., Krehenwinkel, H., Lim, J. Y., Staniczenko, P., Callaghan, J., Andersen, J. C., ... & Gillespie, R. G. (2023). Ecological network structure in response to community assembly processes over evolutionary time. Molecular Ecology, 32(23), 6489-6506.
  23. Gillespie, R., Bik, H., Hickerson, M., Krehenwinkel, H., Overcast, I., & Rominger, A. (2023). Insights into Ecological & Evolutionary Processes via community metabarcoding. Molecular Ecology.
  24. Cerca, J., Cotoras, D. D., Santander, C. G., Bieker, V. C., Hutchins, L., Morin‐Lagos, J., ... & Gillespie, R. G. (2023). Multiple paths toward repeated phenotypic evolution in the spiny‐leg adaptive radiation (Tetragnatha; Hawai'i). Molecular Ecology, 32(18), 4971-4985.
  25. Huszarik, M., Röder, N., Eberhardt, L., Kennedy, S., Krehenwinkel, H., Schwenk, K., & Entling, M. H. (2023). External DNA contamination and efficiency of bleach decontamination for arthropod diet analysis. Environmental DNA, 5(3), 540-550.
  26. Mayer, M., Böning, P., Lima, A. P., Krehenwinkel, H., Bitar, Y. O., Bernarde, P. S., ... & Lötters, S. (2023). Disentangling the biogeographic history of a truly pan-Amazonian amphibian–the case of the three-striped poison frog, Ameerega trivittata (Dendrobatidae: Colostethinae). Salamandra, 59(1).

  27. Weber, S., Junk, I., Brink, L., Wörner, M., Künzel, S., Veith, M., Teubner, D., Klein, R., Paulus, M. and Krehenwinkel, H. (2023). Molecular diet analysis in mussels and other metazoan filter feeders and an assessment of their utility as natural eDNA samplers. Molecular Ecology Resources.

  28. Krehenwinkel, H., Weber, S., Broekmann, R., Melcher, A., Wolf, R., Hochkirch, A., Kennedy, S.R., Koschorrek, J., Künzel, S., Müller, C. and Reztlaff, R., (2022). Environmental DNA from archived leaves reveals widespread temporal turnover and biotic homogenization in forest arthropod communities. Elife.

  29. Krehenwinkel, H., Weber, S., Künzel, S. and Kennedy, S.R., 2022. The bug in a teacup—monitoring arthropod–plant associations with environmental DNA from dried plant material. Biology Letters, 18(6), p.20220091.

  30. Arribas, P., Andújar, C., Bohmann, K., DeWaard, J.R., Economo, E.P., Elbrecht, V., Geisen, S., Goberna, M., Krehenwinkel, H., Novotny, V. and Zinger, L., 2022. Toward global integration of biodiversity big data: a harmonized metabarcode data generation module for terrestrial arthropods. GigaScience, 11.

  31. Pomerantz, A., Sahlin, K., Vasiljevic, N., Seah, A., Lim, M., Humble, E., Kennedy, S., Krehenwinkel, H., Winter, S., Ogden, R. and Prost, S., 2022. Rapid in situ identification of biological specimens via DNA amplicon sequencing using miniaturized laboratory equipment. Nature Protocols, pp.1-29.

  32. Kennedy, S. R., Lim, J. Y., Adams, S. A., Krehenwinkel, H., & Gillespie, R. G. (2022). What is adaptive radiation? Many manifestations of the phenomenon in an iconic lineage of Hawaiian spiders. Molecular Phylogenetics and Evolution, 175, 107564.

  33. Fliedner, A., Rüdel, H., Göckener, B., Krehenwinkel, H., Paulus, M. and Koschorreck, J., 2022. Environmental specimen banks and the European Green Deal. Science of The Total Environment, 852, p.158430.

  34. Kennedy, S., Calaor, J., Zurápiti, Y., Hans, J., Yoshimura, M., Choo, J., Andersen, J.C., Callaghan, J., Roderick, G.K., Krehenwinkel, H. and Rogers, H., 2022. Richness and resilience in the Pacific: DNA metabarcoding enables parallelized evaluation of biogeographic patterns. Molecular Ecology.

  35. Overcast, I., Achaz, G., Aguilée, R., Andújar, C., Arribas, P., Creedy, T.J., Economo, E.P., Etienne, R.S., Gillespie, R., Jacquet, C. and Jay, F. (2022): Towards a genetic theory of island biogeography: Inferring processes from multidimensional community‐scale data. Global Ecology and Biogeography.

  36. Armstrong, E.E., Perez‐Lamarque, B., Bi, K., Chen, C., Becking, L.E., Lim, J.Y., Linderoth, T., Krehenwinkel, H. and Gillespie, R.G., 2022. A holobiont view of island biogeography: Unravelling patterns driving the nascent diversification of a Hawaiian spider and its microbial associates. Molecular Ecology, 31(4), pp.1299-1316.

  37. Emerson, B.C., Borges, P.A., Cardoso, P., Convey, P., deWaard, J.R., Economo, E.P., Gillespie, R.G., Kennedy, S., Krehenwinkel, H., Meier, R. and Roderick, G.K., 2022. Collective and harmonised high throughput barcoding of insular arthropod biodiversity: toward a Genomic Observatories Network for islands. Molecular Ecology.

  38. Lim, J.Y., Patiño, J., Noriyuki, S., Cayetano, L., Gillespie, R.G. and Krehenwinkel, H., 2022. Semi‐quantitative metabarcoding reveals how climate shapes arthropod community assembly along elevation gradients on Hawaii Island. Molecular Ecology, 31(5), pp.1416-1429.

  39. Perez-Lamarque, B., Krehenwinkel, H., Gillespie, R. G., & Morlon, H. (2022). Limited Evidence for Microbial Transmission in the Phylosymbiosis between Hawaiian Spiders and Their Microbiota. Msystems, 7(1), e01104-21.

  40. Overcast, I., Ruffley, M., Rosindell, J., Harmon, L., Borges, P., Emerson, B., Etienne, R., Gillespie, R., Krehenwinkel, H., Mahler, D. L., Massol, F., Parent, C., Patiño, J., Peter, B., Week, B., Wagner, C., Hickerson, M., & Rominger, A. (2021). A unified model of species abundance, genetic diversity, and functional diversity reveals the mechanisms structuring ecological communities. Molecular Ecology Resources.

  41. Graham, N. R., Gillespie, R. G., & Krehenwinkel, H. (2021). Towards eradicating the nuisance of NUMTs and noise in molecular biodiversity assessment. Molecular Ecology Resources.

  42. Foley, S., Krehenwinkel, H., Cheng, D. Q., & Piel, W. H. (2021). Phylogenomic analyses reveal a Gondwanan origin and repeated out of India colonizations into Asia by tarantulas (Araneae: Theraphosidae). PeerJ.

  43. Löbbert, A., Schanzer, S., Krehenwinkel, H., Bracher, F., & Müller, C. (2021). Determination of multi pesticide residues in leaf and needle samples using a modified QuEChERS approach and gas chromatography-tandem mass spectrometry. Analytical Methods. 

  44. Sheffer, M. M., Hoppe, A., Krehenwinkel, H., Uhl, G., Kuss, A. W., Jensen, L., Jensen, C., Gillespie, R. G., Hoff, K. J., & Prost, S. (2021). Chromosome-level reference genome of the European wasp spider Argiope bruennichi: A resource for studies on range expansion and evolutionary adaptation. Gigascience.

  45. Arribas, P., Andujar, C., Bidartondo, M., Bohmann, K., Coissac, E., Creer, S., deWaard, J., Elbrecht, V., Ficetola, G. F., Goberna, M., Kennedy, S., Krehenwinkel, H., Leese, F., Novotny, V., Ronquist, F., Yu, D. W., Zinger, L., Creedy, T., Meramveliotakis, E., Noguerales, V., Overcast, I., Morlon, H., Vogler, A., Papadopoulou​, A., & Emerson, B. (2021). Connecting high-throughput biodiversity inventories – Opportunities for a site-based genomic framework for global integration and synthesis. Molecular Ecology.

  46. Rößler, D. C., Lötters, S., Veith, M., Fugmann, M., Peters, C., Künzel, S., & Krehenwinkel, H. (2020). An amplicon sequencing protocol for attacker identification from DNA traces left on artificial prey. Methods in Ecology and Evolution11(10), 1338-1347.

  47. Kennedy, S. R., & Krehenwinkel, H. (2020). DNA barcoding and community assembly—A simple solution to a complex problem. Molecular Ecology.

  48. Kennedy, S. R., Tsau, S., Gillespie, R., & Krehenwinkel, H. (2020). Are you what you eat? A highly transient and prey‐influenced gut microbiome in the grey house spider Badumna longinqua. Molecular Ecology.

  49. Kennedy, S. R., Prost, S., Overcast, I., Rominger, A. J., Gillespie, R. G., & Krehenwinkel, H. (2020). High-throughput sequencing for community analysis: the promise of DNA barcoding to uncover diversity, relatedness, abundances and interactions in spider communities. Development Genes and Evolution, 1-17.

  50. de Kerdrel, G. A., Andersen, J. C., Kennedy, S. R., Gillespie, R., & Krehenwinkel, H. (2020). Rapid and cost-effective generation of single specimen multilocus barcoding data from whole arthropod communities by multiple levels of multiplexing. Scientific Reports, 10(1), 1-12.

  51. Krehenwinkel, H., Pomerantz, A., & Prost, S. (2019). Genetic Biomonitoring and Biodiversity Assessment Using Portable Sequencing Technologies: Current Uses and Future Directions. Genes, 10(11), 858.

  52. Rößler, D. C., Ogan, S., Curio, E., & Krehenwinkel, H. (2019). Ability makes a thief: vision, learning, and swift escape help kleptoparasitic hover wasps not to fall prey to their spider hosts. Behavioral Ecology and Sociobiology, 73(11), 152.

  53. Krehenwinkel, H., Meese S., Mayer, C., Ruch, J., Schneider, J., Bilde T., Künzel S., Henderson J., Russack J., Simison, W.B., Gillespie R.G., Uhl G. (2019) A guide to cost efficient microsatellite isolation and genotyping by next generation sequencing. Journal of Arachnology.

  54. Krehenwinkel, H., Pomerantz, A., Henderson, J. B., Kennedy, S. R., Lim, J. Y., Swamy, V., ... & Prost, S. (2019). Nanopore sequencing of long ribosomal DNA amplicons enables portable and simple biodiversity assessments with high phylogenetic resolution across broad taxonomic scale. GigaScience, 8(5), giz006.

  55. Krehenwinkel, H., Kennedy, S. R., Adams, S. A., Stephenson, G. T., Roy, K., & Gillespie, R. G. (2019). Multiplex PCR targeting lineage specific SNP s‐A highly efficient and simple approach to block out predator sequences in molecular gut content analysis. Methods in Ecology and Evolution.

  56. Kennedy, S., Lim, J. Y., Clavel, J., Krehenwinkel, H., & Gillespie, R. G. (2019). Spider webs, stable isotopes and molecular gut content analysis: Multiple lines of evidence support trophic niche differentiation in a community of Hawaiian spiders. Functional Ecology.

  57. Andersen, J. C., Oboyski, P., Davies, N., Charlat, S., Ewing, C., Meyer, C., ... & Gillespie, R. G. (2019). Categorization of species as native or non‐native using DNA sequence signatures without a complete reference library. Ecological Applications, e01914.

  58. Foley, S., Lüddecke, T., Cheng, D. Q., Krehenwinkel, H., Künzel, S., Longhorn, S. J., ... & Piel, W. H. (2019). Tarantula phylogenomics: A robust phylogeny of deep theraphosid clades inferred from transcriptome data sheds light on the prickly issue of urticating setae evolution. Molecular Phylogenetics and Evolution, 140, 106573.

  59. Bauer, T., Feldmeier, S., Krehenwinkel, H., Wieczorrek, C., Reiser, N., & Breitling, R. (2019). Steatoda nobilis, a false widow on the rise: a synthesis of past and current distribution trends. NeoBiota, 42, 19.

  60. Schmidt, J. M., Whitehouse, T. S., Green, K., Krehenwinkel, H., Schmidt-Jeffris, R., & Sial, A. A. (2019). Local and landscape-scale heterogeneity shape spotted wing drosophila (Drosophila suzukii) activity and natural enemy abundance: Implications for trophic interactions. Agriculture, Ecosystems & Environment, 272, 86-94.

  61. Krehenwinkel, H., Kennedy, S. R., Rueda, A., Lam, A., & Gillespie, R. G. (2018): Scaling up DNA barcoding–Primer sets for simple and cost-efficient arthropod systematics by multiplex PCR and Illumina amplicon sequencing. Methods in Ecology and Evolution.

  62. Lüddecke T., Krehenwinkel, Canning G., Glaw F., Longhorn SJ, Tänzler R., Ingo Wendt I., Vences M. (2018): Discovering the silk road: Nuclear and mitochondrial sequence data resolve the evolutionary history of theraphosid spiders (Araneae: Mygalomorphae). Molecular phylogenetics and Evolution.

  63. Krehenwinkel H., Fong M., Huang E.G., Kennedy S.R., Suzuki N., Cayetano L., Gillespie R. (2018): DNA degradation bias in metabarcoding studies of arthropod communities and their associated microbiota collected by passive sampling devices. PlosOne.

  64. Krehenwinkel H., Wolf M., Lim J.Y., Rominger A.J., Simsion W.B., Gillespie R. (2017): Estimating and mitigating amplification bias in qualitative and quantitative arthropod metabarcoding. Scientific Reports.

  65. Wawer W., Rutkowski R., Krehenwinkel, Lutyk D., Pusz-Bocheńska K., Bogdanowicz. W. (2017): Population stability of the expansive wasp spider (Argiope bruennichi) at the edge of the range. Journal of Arachnology.

  66. Krehenwinkel H., Kennedy S., Pekár S. & Gillespie R. (2017). A cost efficient and simple protocol to enrich prey DNA from extractions of predatory arthropods for large scale gut content analysis by Illumina sequencing. Methods in Ecology and Evolution, 8, 126-134.

  67. Krehenwinkel H., Rödder D., Năpăruş–Aljančič M., Kuntner M. (2016): Rapid genetic, morphological and ecological differentiation during the northern range expansion of the venomous yellow sac spider Cheiracanthium punctorium in Europe. Evolutionary Applications, 9, 1229-1240.

  68. Freudenschuss, M., Grabolle A., Krehenwinkel H. (2016): Gambaquezonia curioi – a new species of Gambaquezonia from the Philippine island Panay (Araneae: Salticidae). Arachnology, 17, 25-27.

  69. Krehenwinkel H., Graze M., Rödder D., Tanaka K., Baba Y., Muster C., Uhl G. (2016): A phylogeographic survey of a highly dispersive spider reveals Eastern Asia as a major glacial refugium for Palearctic fauna. Journal of Biogeography, 43, 1583–1594.

  70. Krehenwinkel H., Rödder D., Tautz D. (2015): Eco‐Genomic analysis of the poleward range expansion of the wasp spider Argiope bruennichi shows rapid adaptation and genomic admixture. Global change biology, 21, 4320–4332. * Research highlight in Nature Climate Change

  71. Krehenwinkel H., Pekar S. (2015): An analysis of factors affecting genotyping success from museum specimens reveals an increase of genetic and morphological variation during a historical range expansion of a European spider. PloS one, 10, e0136337.

  72. Jäger P., Li S., Krehenwinkel H. (2015): Morphological and molecular taxonomic analysis of Pseudopoda Jäger, 2000 (Araneae: Sparassidae: Heteropodinae) in Sichuan Province, China. Zootaxa, 3999, 363–392.

  73. Jäger P., Krehenwinkel H. (2015): May gen. n. (Araneae: Sparassidae): a unique lineage from southern Africa supported by morphological and molecular features. African Invertebrates, 56, 365–392.

  74. Zimmer S.M., Krehenwinkel H., Schneider J. (2014): Rapid range expansion is not restricted by inbreeding in a sexually cannibalistic spider. PlosOne, 7, e95963.

  75. Krehenwinkel H., Tautz D. (2013): Northern range expansion of European populations of the wasp spider Argiope bruennichi is associated with global–warming–correlated genetic admixture and population specific temperature adaptations. Molecular Ecology 22, 2232–2248.

  76. Krehenwinkel H., Curio E., Tacud J., Haupt J. (2009): On Telyphonoidespanayensis  gen. et sp. nov. (Arachnida: Uropygi: Telyphonidae), a new genus and a new species of whip scorpions from Panay Island (Philippines). Arthropoda Selecta 18, 139–143.

    Books & Book chapters

  • Krehenwinkel H. et al. (2023): Metabarcoding for Biodiversity Estimation Encyclopedia of Biodiversity

  • Krehenwinkel H., Maerklin T., Kroes T. (2008): Ornamentvogelspinnen– die Gattung Poecilotheria.  Herpeton Verlag, Offenbach.

    Non peer reviewed publications

  1. Leese, F., Woppowa, L., Bálint, M., Höss, S., Krehenwinkel, H., Lötters, S., ... & Züghart, W. (2023). DNA-basierte Biodiversitätsanalysen im Natur-und Umweltschutz: Welche Optionen haben wir für eine Standardisierung? Eine Handlungsempfehlung aus Forschung und Praxis.
  2. Melcher, A. C., Weber, S., Calaor, J., Kennedy, S., Gajski, D., Hörrmann, T., ... & Krehenwinkel, H. (2024). Spiders as natural DNA samplers can recover arthropod community diversity and biotic interactions across space and time.

  3. Overcast, I., Calderon-Sanou, I., Creer, S., Dominguez-Garcia, V., Hagen, O., Hickerson, M., ... & Gillespie, R. G. (2025). The distribution of genetic diversity in ecological communities: A unifying measure for monitoring biodiversity change.

  4. Sheffer, M.M., Zander, L., Schulze, B., Mouginot, P., Naef, T., Kreyling, J., Gillespie, R.G., Hoff, K.J., Prost, S., Krehenwinkel, H. and Uhl, G., 2021. Rapid ecological and evolutionary divergence during a poleward range expansion. bioRxiv.

  5. Rominger A.J., Overcast I. Krehenwinkel H., Gillespie R., Harte J., Hickerson M.J. (2017): Linking evolutionary and ecological theory illuminates non-stationary biodiversity. arXiv preprint arXiv:1705.04725. 

  6. Krehenwinkel H., Morra M. (2015): Erhaltungszucht und Wiederauswilderung von Vogelspinnen – eine kritische Analyse. Arachne 20:4–15.

  7. Krehenwinkel H. (2015): Forschung ist der Schlüssel zum Artenschutz – ein DeArGe–Stipendium für Arachnologen. Arachne 20:24–26.​​​​​​​

  8. Jaeger P., Krehenwinkel H. (2012): Wespenspinnen und der Wert wissenschaftlicher Sammlungen. Natur Forschung Museum 142: 94–101.​​​​​​​

  9. Krehenwinkel H. (2008): Eine neue, baumbewohnende Orphnaecus–Art von der Philippinen–Insel Panay. Arachne13: 23–36.